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The code base is the main difference, but there's a difference in philosophy borne out by the code as well. MUSHes tend to focus on the cooperative storytelling part, and things like combat or spell results are often decided beforehand cooperatively. On a MUD, things like combat/spells are a function of the code.
MUD stands for Multi-User Domain/Dungeon, while MuX stands for Multi-User eXperience. MUSH stands for Multi-User Shared Hallucination. MURPE stands for Multi-User Roleplaying Experience, and sometimes gets slapped on the RP-enforced MUDs.
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