My answer: Be ready for both. Players may or may not run with the storyline. It may drop into a lull. They may go off into a dead-end tangent.
The plot should certainly be flexible enough to allow the players to affect where it goes over time, but you should also have a plan for providing guidance and a nudge here and there when required.
It can look like you're winging it, even if you've got a net.
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